D4.2.3 EQUILIBRIUM AND EVOLUTIONARY CHANGE
πDefinition Table
| Term | Definition |
|---|---|
| Hardy-Weinberg equilibrium | Condition where allele and genotype frequencies remain constant in a population. |
| Evolutionary change | Alteration of allele frequencies across generations. |
| Punctuated equilibrium | Long periods of stasis interrupted by rapid evolutionary events. |
| Gradualism | Slow, steady accumulation of evolutionary changes. |
| Stabilising selection | Selection favouring intermediate phenotypes. |
| Directional selection | Selection favouring one extreme phenotype. |
πIntroduction
Populations are not static: they evolve under selective pressures, genetic drift, gene flow, and mutation. However, in the absence of these forces, populations remain in Hardy-Weinberg equilibrium. Evolutionary change can be gradual or punctuated, and selection can stabilise, diversify, or direct populations towards new adaptive peaks. These patterns provide a framework for understanding both microevolutionary changes and macroevolutionary trends in the fossil record
π Hardy-Weinberg Equilibrium
- States that allele frequencies remain constant if no evolutionary forces act.
- Assumptions: large population, random mating, no mutation, no migration, no selection.
- Provides a null model for studying evolution.
- Deviations reveal which evolutionary processes are at work.
- Useful in population genetics, epidemiology, and conservation biology.

π§ Examiner Tip: Show ability to apply Hardy-Weinberg calculations. Simply defining it without calculation limits marks.
π Types of Natural Selection
- Stabilising selection: reduces variation, favours intermediate traits.
- Directional selection: shifts population towards one extreme (e.g., antibiotic resistance).
- Disruptive selection: favours both extremes, may lead to speciation.
- Balancing selection maintains multiple alleles (e.g., sickle-cell trait).
- Selection modes shape population equilibrium and change.
𧬠IA Tips & Guidance: Students could collect data on variation in a measurable trait (e.g., seed size) and test if observed distribution fits stabilising or directional selection.
π Gradualism vs Punctuated Equilibrium

- Gradualism: Darwinβs view β small changes accumulate over long time.
- Punctuated equilibrium: Eldredge & Gould β long stasis interrupted by rapid speciation.
- Fossil record supports both: some lineages show stasis, others rapid bursts.
- Environmental shifts (mass extinctions, climate change) often trigger punctuation.
- Together, they reflect multiple evolutionary tempos.
π EE Focus: An EE could examine fossil evidence for tempo of evolution, e.g., comparing trilobite stasis vs mammalian radiation post-dinosaur extinction
π Microevolution and Macroevolution
- Microevolution: allele frequency shifts within populations.
- Macroevolution: large-scale patterns (speciation, extinction).
- Same processes drive both, but at different scales.
- Links population genetics with paleobiology.
- Demonstrates continuity of evolutionary theory.
β€οΈ CAS Link: Students could create interactive models showing Hardy-Weinberg dynamics or demonstrate selection types with real-life analogies for school outreach.
π Real-World Connection: Hardy-Weinberg used in medical genetics (tracking allele frequencies in diseases), conservation (managing endangered species), and agriculture (maintaining crop diversity).
π Long-Term Evolutionary Dynamics
- Evolution is not linear but shaped by interactions of drift, selection, flow, mutation.
- Extinctions and radiations punctuate long-term patterns.
- Coevolution drives reciprocal adaptations.
- Evolutionary arms races explain rapid trait changes (predator-prey, host-pathogen).
- Stability and change are complementary features of evolution.
π TOK Perspective: Models like Hardy-Weinberg and punctuated equilibrium simplify reality. TOK issue: How far can simplified models capture complex natural processes without distorting understanding?